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MI5 security officer Eve Polastri is bored in her job and yearns for a more exciting life. When a Russian politician is murdered, Eve is tasked with protecting the only witness and soon finds herself on a collision course with a violent and extraordinary assassin, Villanelle.
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In the aftermath of her recent assignment, Villanelle is ordered to take a break. Never one to do as she's told, she goes ahead with her next mission regardless. Meanwhile, Eve is given a dream opportunity to join a secret MI6 unit tracking Villanelle and the shady organization she works for.
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When Villanelle kills a Chinese colonel at a kink clinic in Berlin, Eve and Bill travel out to investigate. While Eve and Bill chase up a number of promising leads, Villanelle enjoys the cat-and-mouse of their proximity. But as they draw closer, Eve comes to realize that this is much more than a game.
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Intel from Berlin points to the existence of a mole, prompting Eve to undertake her first surveillance operation. Meanwhile, Villanelle is sent to England to eliminate a member of British Intelligence. Villanelle wonders, could it be Eve? Meanwhile Eve, in emotional turmoil, now knows how high the stakes are.
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Having survived a terrifying close call with Villanelle, Eve now has the mole ensconced in a safe house and is buzzing. They have an exciting opportunity to glean information and crack this thing open.
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Konstantin tells Villanelle that Nadia did not die as she claims but is in a Moscow Jail. The team also finds out that Nadia is still alive in a Moscow jail. Carolyn uses a past connection which happens to be one of the handlers ,Konstantin. As Eve and Carolyn speak with Nadia, she at first is willing to speak about Villanelle but Konstantin shows up and Nadia clams up. Meanwhile Villanelle has also gotten herself in the same Moscow Jail with the intention of Konstantin getting her out after she takes care of Nadia. Eve asks Kenny to look into any and all correspondence between his mother Carolyn and the Russians as Eve fears the worst is about to happen.
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Kenny is now in Moscow, as Carolyn comes home from an allnighter and tells Eve and Kenny to go back to London as Nadia has been killed while waiting to be released to the UK Authorities. Eve and Carolyn go to the hotel restaurant and see Konstantin, who like Carolyn is still wearing the clothes they wore yesterday. Back in prison they give a roommate to Villanelle while she is in solitary who tries to kill her but fails as Villanelle kills her killer. The guards rush her out to transfer her to another location which is to escape. She learns of a new handler called Anton who instructs her to kill Konstantin as she turns and kills Anton. Back in the makeshift office, Kenny gives Eve love letters from Konstantin and his mother, Carolyn. More evidence turns up as Kenny and Eve defy Carolyn's order to return to London while Konstantin is confronted by Villanelle, as he is her next target. Kenny breaks into the cctv at the prison to his and Eve's horror, soon to discover explosive findings.
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Villanelle's mission is proving anything but straightforward, forcing her to take drastic measures. Meanwhile, Eve makes a shocking discovery, which leads her right to Villanelle and an electrifying confrontation from which neither will emerge unscathed.
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Eve is reeling from her act of violence against Villanelle and doesn't know whether she is alive or dead. A paranoid and severely injured Villanelle manages to get herself to the hospital for treatment. Carolyn approaches Eve with an opportunity.
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After Eve is shot by Villanelle, she's trying to move on. Villanelle is also moving on, until she is approached by someone from her past. Carolyn is being undermined at work and Kenny is looking for more information about the twelve.
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Eve and Carolyn are reeling. Eve strikes up an unlikely alliance with Kenny's colleagues at the Bitter Pill. Villanelle finds that management isn't all it's cracked up to be.
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Villanelle agrees to a job which will take her back to London; Carolyn's Moscow contacts come in use when Eve and the Bitter Pill team unravel a potential lead to The Twelve; Konstantin finds himself under pressure from all sides.
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Eve is trying to patch things up with Niko; Villanelle receives good news about her upcoming promotion but can't shake the feeling there's still something missing from her life; Dasha seeks a solution to Villanelle's erratic behavior.
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Villanelle visits her family in Russia, and while the quirky reunion starts as an opportunity to reconnect and answer unknowns about her childhood, a profound moment of clarity sends her on a disturbing and downward spiral.
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As the investigation continues, Carolyn finds out that information is being withheld by those close to her. Eve finds a lead which might bring her closer to Villanelle, but Villanelle has had enough and decides on a different path.
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Lack of progress in the investigation is frustrating Caroline. Eve searches for a clue to her Villanelle's whereabouts. Villanelle and Dasha team together for a kill. Konstantin is crumbling under pressure.
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